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Paréntesis

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Palabra sobre la que consulta: 
Paréntesis
Consulta: 

¿Entre dos paréntesis seguidos debo colocar punto y coma, coma o punto según la información contenida en cada uno? Por ejemplo: Particularmente variables son los mutualismos de defensa mediados por nectarios extraflorales (plantas mirmecófilas) (Boecklen 1984).

Respuesta: 

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